Step 1: The tapetum is the innermost layer of the microsporangium (anther wall) and consists of cells with dense cytoplasm and generally more than one nucleus.
Step 2: Its primary function is to provide nutrition to the developing microspores (pollen grains).
Step 3: It also secretes various enzymes, hormones, and the fatty substance sporopollenin, which forms the tough outer layer (exine) of the pollen grain.