Step 1: Understanding the Concept:
The australopithecines are a group of extinct hominins. Paleoanthropologists classify them into two main morphological groups: "gracile" and "robust." This classification is based primarily on features related to chewing, such as the size of the teeth, jaws, and chewing muscles.
- Gracile australopithecines had a lighter build, especially in the skull and teeth. Their jaws were less massive, and their molars were smaller.
- Robust australopithecines (often placed in the genus Paranthropus) had massive jaws, enormous molars, and prominent sagittal crests on their skulls for the attachment of large chewing muscles.
Step 2: Detailed Explanation:
Let's classify each species:
- A. A. africanus: This is the classic example of a gracile australopithecine from South Africa.
- B. A. boisei: Also known as Paranthropus boisei, this species is the epitome of a "hyper-robust" australopithecine from East Africa, known for its massive jaws and molars. It is not gracile.
- C. A. afarensis: This East African species, famously represented by the "Lucy" fossil, is a gracile australopithecine.
- D. A. anamensis: This is one of the earliest australopithecine species and is considered a gracile form.
Step 3: Final Answer:
The species classified as gracile australopithecines are A. africanus, A. afarensis, and A. anamensis. A. boisei is a robust australopithecine. Therefore, the correct combination is A, C, and D only.