Phloem consists of several types of cells including sieve elements, companion cells, phloem parenchyma, and fibers.
Among these, phloem sieve elements are unique because they are living at maturity but lack a nucleus.
This structural adaptation allows for more efficient translocation of sugars and other organic substances throughout the plant.
However, the loss of the nucleus also makes them dependent on adjacent companion cells, which provide metabolic support.
Other phloem cells like phloem parenchyma and companion cells retain their nuclei and perform different functions.