Step 1: Understand the successional pattern.
Ecological succession often begins with early colonisers that are excellent at dispersal and rapid establishment.
Over time, these species are replaced by late-successional species that are superior competitors for local resources.
Step 2: Analyze option (A).
In the early stages of succession, resources such as space, light, and nutrients are usually abundant, not limiting.
Resource limitation typically increases later as biomass accumulates.
Hence, (A) is incorrect.
Step 3: Analyze option (B).
This statement relates to keystone species and trophic facilitation, not directly to the coloniser–competitor replacement pattern.
Hence, (B) is incorrect.
Step 4: Analyze option (C).
Trees, which often dominate climax terrestrial communities, generally have high competitive ability but relatively low dispersal ability.
The statement reverses these traits.
Hence, (C) is incorrect.
Step 5: Analyze option (D).
A well-established ecological principle is the tradeoff between dispersal ability and competitive ability.
Good colonisers disperse widely but are weak competitors, while late-successional species disperse poorly but compete effectively.
This tradeoff directly explains the observed successional pattern.
Hence, (D) is correct.
Step 6: Conclusion.
The replacement of good colonisers by good competitors during succession is best explained by:
\[
\boxed{(D)}
\]